Canh M. Nguyen * , Van T. M. Nguyen , Dung A. Pham , Hai T. M. Nguyen , Vu T. Cong , & Khoi M. Ka

* Correspondence: Nguyen Minh Canh (email: nmcanh@hcmuaf.edu.vn)

Main Article Content

Abstract

Poor evergreen broadleaf natural forests constitute an important component of tropical forest ecosystems, playing a critical role in maintaining biodiversity and providing a foundation for sustainable forest restoration. This study aimed to assess stand structure, natural regeneration characteristics, and tree species diversity of poor evergreen broadleaf natural forests on lowland hills in Ninh Dien, Hoa Hoi, and Phuoc Vinh communes, Tay Ninh Province, as a scientific basis for forest management and restoration. A total of 47 tree species belonging to 24 families were recorded, with Dipterocarpaceae as the dominant family; Fabaceae, Sapindaceae, Chrysobalanaceae, and Malvaceae also contributed substantially to the stand composition. Five species of ecological importance were identified, including Vatica odorata, Hopea ferrea, Nephelium chryseum, Parinari annamensis, and Dipterocarpus costatus. The mean stand density was 516 trees/ ha, with an average growing stock of 84.5 m³/ha. Diameter distribution exhibited a left-skewed unimodal pattern described by the Weibull function, while height distribution followed a normal distribution, indicating an intermediate post-disturbance recovery stage with a relatively simple stand structure. Natural regeneration was abundant, with a density of 4,288 individuals/ha, dominated by seed-origin regeneration and primarily composed of native species. Shannon-Wiener diversity and Pielou evenness indices indicated a relatively balanced species composition without absolute dominance. Based on these findings, the study proposes silvicultural measures, including assisted natural regeneration, enrichment planting with native species, and long-term monitoring of forest dynamics, contributing to the restoration and sustainable management of natural forests in the study area.

Keywords: Biodiversity, Forest structure, Natural forest rehabilitation, Natural regeneration, Poor evergreen broad-leaved forest

Article Details

References

Bui, T. H. (2024). Structural characteristics of recovering natural forests at states IIIA1 and IIIA2 in Ben En National Park, Thanh Hoa Province. Hong Duc University Journal of Science 59, 75-83. https://hdujs.edu.vn/index. php/hdujs/article/view/90.

Cao, T. T. H., & Do, H. H. (2019). Structural characteristics and tree species diversity of natural forest at state IIIA1 on the eastern and western slopes of the ecological restoration zone of Ba Vi National Park, Hanoi. Journal of Forestry Science and Technology 1, 65-74.

Chazdon, R. L., Norden, N., Colwell, R. K., & Chao, A. (2022). Monitoring recovery of tree diversity during tropical forest restoration: Lessons from long-term trajectories of natural regeneration. Philosophical Transactions of The Royal Society B 378, 20210069. https://doi.org/10.1098/ rstb.2021.0069.

Dawkins, H. C. (1958). The management of natural tropical high-forest with special reference to Uganda. Oxford, England: Commonwealth Forestry Institute, University of Oxford.

Do, H. H., Cao, T. T. H., & Nguyen, T. H. (2023). Some characteristics of natural regeneration in three forest states in Vu Quang National Park, Ha Tinh Province. Journal of Forestry Science and Technology (4), 28-36. https://journal.vnuf.edu.vn/vi/article/view/649.

FAO (Food and Agriculture Organization of the United Nations). 2018. The state of the world’s forests 2018 - Forest pathways to sustainable development. Rome, Italy: Food and Agriculture Organization of the United Nations.

Fernando, E. (1998). Forest formations and flora of the Philippines: Handout in FBS 21. College of Forestry and Natural Resources, University of the Philippines at Los Banos, Laguna, Philippines.

GSPV (Government of the Socialist Republic of Viet Nam). (2021). Decree No. 84/2021/NDCP dated September 22, 2021, amending and supplementing a number of articles of Decree No. 06/2019/ND-CP on the management of endangered, precious, and rare forest flora and fauna and the implementation of the Convention on International Trade in Endangered Species of Wild Fauna and Flora (CITES). Retrieved April 12, 2025, from https:// thuvienphapluat.vn/van-ban/Tai-nguyen-Moitruong/Nghi-dinh-84-2021-ND-CP-sua-doiNghi-dinh-06-2019-ND-CP-quan-ly-thuc-vatrung-488788.aspx.

Hernández-Ruedas, M. A., Arroyo-Rodríguez, V., Meave, J. A., Martínez-Ramos, M., IbarraManríquez, G., Martínez, E., Jamangapé, G., Melo, F. P. L., & Santos, B. A. (2014). Conserving tropical tree diversity and forest structure: The value of small rainforest patches in moderatelymanaged landscapes. Plos One 9(6), e98931. https://doi.org/10.1371/journal.pone.0098931.

Le, D. T., Tran, X. T., Le, T. H., & Le, A. T. (2023). Structural characteristics and tree species diversity of evergreen closed forest in Muong Phang, Dien Bien Province. Journal of Forestry Science and Technology 12(5), 96-106. https://journal.vnuf.edu.vn/vi/article/view/116.

Liu, J. (2025). Progress in research on the effects of environmental factors on natural forest regeneration. Frontiers in Forests and Global Change 8, 1525461. https://doi.org/10.3389/ ffgc.2025.1525461.

MARD (Ministry of Agriculture and Rural Development). (2022). Circular No. 17/2022/ TT-BNNPTNT dated October 27, 2022, amending and supplementing a number of articles of Circular No. 29/2018/TT-BNNPTNT on silvicultural measures. Retrieved April 12, 2025, from https://thuvienphapluat.vn/ van-ban/Tai-nguyen-Moi-truong/Thongtu-17-2022-TT-BNNPTNT-sua-doi-Thongtu-29-2018-TT-BNNPTNT-bien-phap-lamsinh-535557.aspx.

MARD (Ministry of Agriculture and Rural Development). (2018). Circular No. 28/2018/ TT-BNNPTNT dated November 16, 2018, on sustainable forest management. Retrieved April 12, 2025, from https://thuvienphapluat. vn/van-ban/Tai-nguyen-Moi-truong/Thongtu-28-2018-TT-BNNPTNT-quy-dinh-ve-quanly-rung-ben-vung-402853.aspx.

Margalef, R. (1968). Perspectives in ecological theory. Chicago, USA: University of Chicago Press.

NASRV (National Assembly of the Socialist Republic of Viet Nam). (2017). Law No. 16/2017/QH14 dated November 15, 2017. Law on Forestry. Retrieved August 12, 2025, from https:// thuvienphapluat.vn/van-ban/Linh-vuc-khac/ Luat-lam-nghiep-367277.aspx.

Ngo, T. P., Phan, M. X., & Nguyen, M. C. (2023). Structural characteristics and tree species diversity of evergreen broad-leaved forests in Phuoc Binh National Park, Ninh Thuan Province. Journal of Agriculture and Rural Development 1, 71–81.

Nguyen, H. H., Pham, V. D., Bui, T. D., Trinh, H. M., Nguyen, V. Q., & Nguyen, T. T. (2024). Effects of species and tree size diversity on above-ground biomass in diverse heterogeneous tropical evergreen forests, Quang Binh Province, north central Vietnam. Annals of Forest Research 67(1), 31-40. https://doi.org/10.15287/afr.2024.3293.

Nguyen, M. C., To, Q., Mac, V. C., Nguyen, T. M. H., Nguyen, T. T., Dang, L. T. L., & Ho, T. T. (2025). Forest structure, tree species diversity, and soil carbon stocks in rich evergreen broadleaf natural forest at Lo Go - Xa Mat National Park, Tay Ninh Province. Vietnam Journal of Forest Sciences 14(7), 1130. https://doi. org/10.70169/VJFS.1130.

Nguyen, T. T. H., & Tran, T. T. H. (2014). Study on some structural characteristics of evergreen broadleaf natural forest in Vu Quang National Park, Ha Tinh Province. Journal of Forestry Science 3, 3408-3416.

Pan, Y., Birdsey, R. A., Fang, J., Houghton, R., Kauppi, P. E., Kurz, W. A., Phillips, O. L., Shvidenko, A., Lewis, S. L., Canadell, J. G., Ciais, P., Jackson, R. B., Pacala, S. W., McGuire, A. D., Piao, S., Rautiainen, A., Sitch, S., & Hayes, D. (2011). A large and persistent carbon sink in the world’s forests. Science 333(6045), 988-993. https://doi. org/10.1126/science.1201609.

PCTNP (People’s Committee of Tay Ninh Province). (2018). Approval of the 2018 land-use plan of Chau Thanh district. Retrieved April 25, 2025, from https://www.tayninh.gov.vn/vi/news/ thong-tin-dat-dai/ph-duy-t-k-ho-ch-s-d-ng--tn-m-2018-c-a-huy-n-ch-u-th-nh-38506.html.

Pielou, E. C. (1975). Ecological diversity. London, England: Wiley - Interscience Publication.

Pretzsch, H. (2009). Forest dynamics, growth and yield: From measurement to model. Berlin, Germany: Springer-Verlag.

Psistaki, K., Tsantopoulos, G., & Paschalidou, A. K. (2024). An overview of the role of forests in climate change mitigation. Sustainability 16(14), 6089. https://doi.org/10.3390/su16146089.

Shannon, C. E., & Weaver, W. (1963). The mathematical theory of communication. Urbana, IL, USA: University of Illinois Press.

Simpson, E. H. (1949). Measurment of diversity. Nature 163(4148), 688. http://dx.doi. org/10.1038/163688a0.

Thai, V. T. (1999). Tropical forest ecosystems in Vietnam. Ha Noi, Vietnam: Science and Technology Publishing House.

Tran, H. (2002). Timber resources of Vietnam. Ha Noi, Vietnam: Agricultural Publishing House.

Vo, D. H. (2015). Carbon stocks in tropical evergreen broadleaf forests in central highlands Vietnam. International Forestry Review 17(1), 20-29. https://www.jstor.org/stable/24310649.

Vo, D. H. (2014). Study on structural characteristics of the upper tree layer of forest state IIA in Yen Lap protection forest, Quang Ninh province. Journal of Forestry Science 3, 3390-3398.

Vo, V. C. (2003). Common plant dictionary. Vol. 1. Ho Chi Minh City, Vietnam: Science and Technology Publishing House.

Vo, V. C. (2004). Common plant dictionary. Vol. 2. Ho Chi Minh City, Vietnam: Science and Technology Publishing House.

Vu, A. D. (2025). Structural characteristics and tree species diversity of poor evergreen broadleaf natural forest on soil mountains in Dau Tieng protection forest, Tay Ninh province (Master’s thesis). Nong Lam University, Ho Chi Minh City, Vietnam.

Williams, B. A., Beyer, H. L., Fagan, M. E., Chazdon, R. L., Schmoeller, M., Sprenkle-Hyppolite, S., Griscom, B. W., Watson, J. E. M., Tedesco, A. M., Gonzalez-Roglich, M., Daldegan, G. A., Bodin, B., Celentano, D., Wilson, S. J., Rhodes, J. R., Alexandre, N. S., Kim, D.-H., Bastos, D., & Crouzeilles, R. (2024). Global potential for natural regeneration in deforested tropical regions. Nature 636(8041), 131-137. https://doi. org/10.1038/s41586-024-08106-4.